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FRANÇAIS                                                        THE PROJECT                                                          ESPAÑOL 
The new classification of Cacti mainly based on molecular data and explained
by Joël Lodé
Volumes 3-4-5-6:
"Description of the Species"

      ±2,000 pages / ±15,000 photos / 190 genera / ±2700 species and subspecies  
updated 25th of May 2024    

A surprise is waiting you at the end of this page!

FORECAST of contents:
- Approximately 2,700 listed taxa (sp. + subsp.) recognised, described and explained in alphabetical order,
- ± 2,000 pages A4 format (+ index of taxa treated),
- ± 1000 photos of new taxa and those omitted in the first two volumes, taken in habitat and in collection,
- + 2,500 photos of seeds of almost all species and subspecies, taken under a digital microscope,
- Complete description, updated for each species and subspecies, and moreover, why they were retained, synonymized or discarded.
- Etymology and eponymy for each species and subspecies,
- Portrait of persons who gave their name to a species or subspecies,
- + 1,000 geographical distribution maps, drastically reduced in number through between others, regroupments of subspecies.
- Description of habitats for each species and subspecies,
- Level of threat for each species and subspecies, presence in cultivation,
- Bibliography,
- Glossary,
- Index and revised synonyms (in a separate booklet).

This is the ambitious project that prefigures the sequel and the end of the first two volumes of the "Taxonomy of the Cactaceae", which presented for the first time in a single work, the genera of Cacti recognised by the author and using both morphological data and molecular phylogenetic studies.

This time it is an even more detailed and complementary huge
research work, description of all taxa recognised by the author (± 2700) and new data, like pictures under digital microscope of the seeds of almost all species and subspecies concerned.

The book is written in English (will be revised and corrected by specialists!), its translation into French will require one more year of work, thus delaying a publication already planned in the long term, considering the amount of work, knowing that the necessary funds for the writing of this work, and its translation do not exist. This work is based on the will and perseverance of the author (and the patience of future readers!), It requires both financial and logistical support; three databases are already available at the bottom of this page: the list of taxa seeds still to be found, as well as the photographs of the taxa not previously included in the first two volumes. If you are a happy man and have seeds or photos wanted for this book, you are welcome to participate in the illustration! The collaborators will of course be cited and thanked with their names in the work, and the illustrations will be duly identified with the copyright of their authors.

digitalisation des graines

The drafting of the book is a long-term task which demands an almost exclusive dedication, which is not remunerated, and even generates expenses. The financial assistance is therefore particularly welcome to carry out this unpublished project which will spread over several years. I would like to thank the Société Française d'Acclimatation  (Society whose aim is to improve the diffusion of the knowledge of gardeners about their climate and the plants of acclimatisation, to favor the diversity of the plantations in gardens, in agreement with the climate) which was the first to value this work and offered a grant for its implementation, also to Dr Alexander Bunkenburg, Alexander Pichler, Philippe Richaud,  Frédéric Carlier, Brice ChéronPeter Quirini, Grzegorz Matuszewski, Marco Alberto Medda, Fabrice ArnaudPhilippe Charbonnier and Al Laius for their financial or material support, as well as Christophe Ludwig for hosting all books data on server, and Jorge Quiñónez as informant.
There is nothing formal, every help is greeted with kindness, and before being embodied in the book with the thanks due to the generous collaborators, it will be pointed out through this page which will honour each step forward towards the final publication of a work necessary to update our knowledge in the field of Cactaceae. In advance, thank you for your involvement.
If you want to help this project to come true, you can also participate with 5, 10, 20 etc. or more Euros, just send the money to the account of cactus-adventures.com on our Paypal account or secure server mentioning (Taxonomy of the Cactaceac Project vol.3-4"*. THANK YOU!
                                                                                           Joël Lodé
* https://www.cactus-aventures.com/secure.php

Started in 2016 for the volumes 3-4, the project is already well under way:

SEEDS under digital microscope:
- Seeds are like fingerprints for cacti; for now,
I have currently taken 8,306 photos of the seeds under digital microscope, which represent about 2,650 taxa. Some come from different sources to verify and ensure their identity; most cacti seed sellers found on the web were solicited, some even offered the seeds, which represent a great help. Many amateurs and specialists as well as botanical gardens and botanists have also participated in this compilation on a voluntary and enthusiastic basis, and all will be thanked and cited in the book

Thanks to many collaborators who will be cited fo their merits in the book, the collection of still missing photos in the first two volumes is well advanced, actually with more than 1,500 photos of new taxa and pictures.

More than 1,000 geographical distribution maps of all taxa (the subspecies would be grouped together with the species on a single map in order to reduce their number) are at the project stage, using the QGIS or DIVA-GIS software for their realization. I still have to find a standard model. For now, most information about this distribution is compiled. Help with the use of this software is welcome!

I have also prepared what will be used for the etymological illustration, the model for the gallery showing the persons who gave their name to a species or a subspecies. It is a very laborious and often unsuccessful research work, many people have not been represented, and there is no portrait or photo of these people known, or more simply I have not managed to find their portrait : for example, the chevalier de Monville, Francisco Ortega respectively for genera Monvillea and Ortegocactus, Richard Grässner for Acanthocephala graessneri, José Picardo for Airampoa picardoi etc. There will be many anecdotes about the lives of these characters, many of which will surprise us.

The text of the descriptions in English is finished (vol. 4), it remains to be revised, but I often go back because of new data that appears, taxonomic changes or new taxa confirmed, e.g. currently on Eriosyce sensu lato.. .  Updated list of genera below. New changes will take place in the list in October 2024.
A long-term work on which I plan to offer a book as complete and updated as possible on Cacti.

Updated list of genera actually treated in Tax. of Cact. vols.3-4

zone de travail

Most recent studies:

2023: a new phylogenetic revision of the genus Cereus by Taylor et al.

2023: a new, very complete phylogenetic revision of the genus Mammillaria revisited by Chincoya et al.

2023: a revision of the genus Leptocereus on Hispaniola by Hoxey et al.

2022: at last, a phylogenetic work on Melocactus by Majure et al.

2022: two revisions of Cacti, from Chile (Walter & Guerrero), and Ecuador (Loaiza).

2022. Grusonia over Corynopuntia in Bárcenas y Hernández, not followed in my work.
2022: a new molecular study on Pilosocereus by Franco-Estrada et al.
2021: Mammillaria made monophyletic! A study by Breslin et al. is in the process of being published in Taxon and shows Mammillaria sensu lato divided into three large groups: Cochemiea sensu lato (including among others the Mammillarias of Baja California as already previously)
anticipated, Coryphantha sensu lato (including Escobaria) and Mammillaria sensu lato (practically the not hooked ones).
September 2020: Lavor et al. carried out a molecular analysis for the genus Pilosocereus with about forty taxa.
June 2020: Barrios
et al. have just published a molecular work on the genus Leptocereus, which shows that Dendrocereus is embedded within a monophyletic clade of Leptocereus.
October 2019: Sánchez et al. lead a third phylogenetic study on the genus Echinocereus, but this time, they give importance to the morphology and included it in their work.
A DNA study is also published on the genus Tephrocactus (M. Laura Las Peñas et al.).

2019: in September, a new phylogenetic work on Cylindropuntia still more complete was done by Majure et al.
Also in August, a complete molecular analysis was published on the genus Epithelantha (Aquino et al.).
2019: in August 2019, a new molecular study (Vázquez-Sánchez et al.) confirmed that Turbinicarpus should be splitted into three genera: Turbinicarpus sensu stricto, Rapicactus and Kadenicarpus, already suggested in Taxonomy of the Cactaceae second volume as the “clade B1”, but expecting more studies.

- I will try to take stock with you by updating this page and this text from time to time according to the progress of the project. Patience is a must!
It is just incredible to see how people are contributing with pictures, seeds, portraits, articles from all over the world: thank you for your help and trust.
Joël Lodé, 2016-2024
contact: joel@cactus-aventures.com

I need your help to make this work the most accurate and as complete as possible: thank you for your contributions!
Wishlist of STILL MISSING SEEDS (updated with only 265 taxa missing of about 2,700!)                        
MISSING PICTURES Wishlist (updated with only 32 species missing)
Wishlist of MISSING PORTRAITS (updated, 155 people missing of about 900!)